Individual based simulations of population dynamics require the availability of growth models with adequate complexity. For this purpose a simple-to-use model (non-linear multiple regression approach) is presented describing somatic growth and reproduction of Daphnia as a function of time, temperature and food quantity. The model showed a good agreement with published observations of somatic growth (r2 = 0.954, n = 88) and egg production (r2 = 0.898, n = 35). Temperature is the main determinant of initial somatic growth and food concentration is the main determinant of maximal body length and clutch size. An individual based simulation was used to demonstrate the simultaneous effects of food and temperature on the population level. Evidently, both temperature and food supply affected the population growth rate but at food concentrations above approximately 0.4 mg Cl−1Scenedesmus acutus temperature appeared as the main determinant of population growth.
Four simulation examples are given to show the wide applicability of the model: (1) analysis of the correlation between population birth rate and somatic growth rate, (2) contribution of egg development time and delayed somatic growth to temperature-effects on population growth, (3) comparison of population birth rate in simulations with constant vs. decreasing size at maturity with declining food concentrations and (4) costs of diel vertical migration. Due to its plausible behaviour over a broad range of temperature (2–20 °C) and food conditions (0.1–4 mg Cl−1) the model can be used as a module for more detailed simulations of Daphnia population dynamics under realistic environmental conditions. 相似文献
The central structure belt in Turpan-Hami basin is composed of the Huoyanshan structure and Qiketai structure formed in late Triassic-early Jurassic, and is characterized by extensional tectonics. The thickness of strata in the hanging wall of the growth fault is obviously larger than that in the footwall, and a deposition center was evolved in the Taibei sag where the hanging wall of the fault is located. In late Jurassic the collision between Lhasa block and Eurasia continent resulted in the transformation of the Turpan-Hami basin from an extensional structure into a compressional structure, and consequently in the tectonic inversion of the central structure belt of the Turpan-Hami basin from the extensional normal fault in the earlier stage to the compressive thrust fault in the later stage. The Tertiary collision between the Indian plate and the Eurasian plate occurred around 55Ma, and this Himalayan orogenic event has played a profound role in shaping the Tianshan area, only the effect of the collision to this area was delayed since it culminated here approximately in late Oligocene-early Miocene. The central structure belt was strongly deformed and thrusted above the ground as a result of this tectonic event. 相似文献
In the metamorphic cores of many orogenic belts, large macroscopic folds in compositional layering also appear to fold one or more pervasive matrix foliations. The latter geometry suggests the folds formed relatively late in the tectonic history, after foliation development. However, microstructural analysis of four examples of such folds suggests this is not the case. The folds formed relatively early in the orogenic history and are the end product of multiple, near orthogonal, overprinting bulk shortening events. Once large macroscopic folds initiate, they may tighten further during successive periods of sub-parallel shortening, folding or reactivation of foliations that develop during intervening periods of near orthogonal shortening. Reactivation of the compositional layering defining the fold limbs causes foliation to be rotated into parallelism with the limbs.Multiple periods of porphyroblast growth accompanied the multiple phases of deformation that postdated the initial development of these folds. Some of these phases of deformation were attended by the development of large numbers of same asymmetry spiral-shaped inclusion trails in porphyroblasts on one limb of the fold and not the other, or larger numbers of opposite asymmetry spirals on the other limb, or similar numbers of the same asymmetry spirals on both limbs. Significantly, the largest disparity in numbers from limb to limb occurred for the first of these cases. For all four regional folds examined, the structural relationships that accompanied these large disparities were identical. In each case the shear sense operating on steeply dipping foliations was opposite to that required to originally develop the fold. Reactivation of the folded compositional layering was not possible for this shear sense. This favoured the development of sites of approximately coaxial shortening early during the deformation history, enhancing microfracture and promoting the growth of porphyroblasts on this limb in comparision to the other. These distributions of inclusion trail geometries from limb to limb cannot be explained by porphyroblast rotation, or folding of pre-existing rotated porphyroblasts within a shear zone, but can be explained by development of the inclusion trails synchronous with successive sub-vertical and sub-horizontal foliations. 相似文献
This paper investigates the role of housing age in constraining residential mobility, measured as the percent of households that have moved into their homes in the past 15 months. The leading explanation for why mobility rates differ so much among regions of the United States has been the overall level of growth. The present analysis shows that the growth effect operates through both the newness of population (migration) and the newness of housing available for occupancy by all local residents. The posited explanation for this housing age effect is that progressively older units contain increasingly settled occupants, yielding fewer opportunities for in-movers in areas with older housing. It is empirically demonstrated that households in older housing have lower likelihood of recent mobility even after controlling for age, tenure, migration status, and state location of residence. The analysis reveals the temporal interdependency of mobility, migration, person age, and housing age. 相似文献
Arid karst landscapes that have been degraded by human activities provide a challenge for rehabilitation and an opportunity
to test ideas about the stability and resilience of limestone ecosystems. The Nullarbor Plain is the largest arid karst area
in Australia (220 000 km2) and is divided into extensive closed karstic depressions separated by low rocky ridges, while the dominant vegetation is
chenopod shrubland. Since European settlement there has been considerable change in the vegetation, with significant reduction
in shrub and grass cover over large areas of the plain. These changes are related to a state and transition model of vegetation
dynamics which incorporates climatic variability, fire history and grazing pressure from sheep, kangaroos and rabbits. A partial
sediment budget using 137Cs inventories reveals local and regional patterns of soil redistribution within this arid karst landscape. Rehabilitation
of eroded soil in pastoral lands has been accomplished at several sites but is labour intensive and vulnerable to climatic
fluctuations. Given the low stock numbers, limited number of people involved, and poor economic returns, it would be sensible
to make pastoral activities on the Nullarbor secondary to conservation priorities. This would necessitate a change in land
ethic to stewardship, with emphasis on rehabilitation and control of feral animals. Management of increased numbers of visitors
to the caves and karst also requires that resource inventories and management plans for each area be drawn up and used.
Received: 1 June 1995 · Accepted: 4 December 1995 相似文献
Influence of water temperature on embryonic and larval development of bream (Abramis brama L.) was stdied. Eggs of bream were incubated at eight constant water temperatures between 13.2 and 26.8°C. The temperature
of 21.1°C gave highest hatching success, with no abnormalities in the eleutheroembryos and lowest mortality observed between
eyed egg stage and the time of hatching. Developmental rate increased with increasing temperature. Duration of embryonic development
(y; hours) decreased with increasing incubation temperature (x; °C) according to the formula: y=910.1−65.88 x+1.318 x2. Larvae were reared at eight constant temperatures ranging from 13.5 to 34.0°C. The instantaneous growth rate in wet weight
increased with increasing test temperature from 13.5 to 29.9°C, and then decreased at higher temperatures. Individual growth
of fish and biomass production rate were highest at 27.9°C. This temperature is considered optimal when food availability
and photoperiod are no limiting factors. 相似文献
Coleps hirtus viridis was the dominant species of the planktonic ciliate community of Lake Fühlinger See (Germany) during the study in 1999 and 2000. Total ciliate densities ranged from 120 to 42,000 ind. l−1 in 1999 and up to 8,000 ind. l−1 in 2000. Coleps contributed up to 98% to both total ciliate abundance and biomass and made up an average of 64% of the total ciliate biomass. Oligotrichs (Rimostrombidium, Strobilidium) dominated the epilimnetic zone, whereas peritrich ciliates (Pelagovorticella, Vorticella) were predominantly located in the hypolimnion. The population maximum of Coleps changed locations from the epilimnion in early summer to the hypolimnion (up to 40,000 ind. l−1) during stratification. High growth rates in the hypolimnion, presence of endosymbiontic algae and the ability to ingest detritus seem to be important for the success.Growth rates of Coleps in June were determined by Landry-Hassett dilution experiments in both the epilimnion and the hypolimnion. The instantaneous growth rates were similar in both layers (0.6 d−1), but a distinctly higher instantaneous mortality was estimated for the epilimnion. These high loss rates may be due to grazing pressure by cladocerans.The significance of the histophagous feeding of Coleps was evaluated through an experiment using killed zooplankton. Parts of Daphnia magna were incorporated at rates of about 1,100 μm3 ind.−1 h−1 by Coleps without endosymbiotic algae and at rates of 500 μm3 ind.−1 h−1 by Coleps with endosymbionts. These high feeding rates support the conclusion that Coleps can use dead organic matter as an additional food source. 相似文献